SNAKE, common name for reptiles in the
order Squamata, which also includes
lizards, and of the suborder Serpentes.
The approximately 2500 different species
known display a great range in
appearance and behavior, from the antics
of the harmless hognose to the stealth of
the deadly mamba. The largest family in
the order, Colubridae, includes the most
common snakes on all continents except
Australia, where the family Elapidae
outnumbers all others. This latter group
includes some of the most poisonous
snakes in the world, such as the cobra and coral snake. Another important
family of
venomous snakes is Viperidae, which includes the pit vipers, such as the
rattlesnake and the fer-de-lance, and the true vipers, such as the adder.
The largest
snakes in the world are members of the family Boidae, which includes the
boa and
the python. Some members of this family never attain a length of more than
0.6 m
(2 ft), but the largest may grow to more than 9 m (30 ft).
Physical Characteristics
The body of the snake is long and usually cylindrical, although the bodies
of sea
snakes and many tree-dwelling snakes are flattened on the sides. The body
is
covered with horny scales arranged in regular rows and usually overlapping,
like tiles
on a roof. The scales on the back and sides are generally smaller than
those on the
top of the head and along the under, or ventral, surface. The scales on
the upper, or
dorsal, surface of the snake may be smooth or may have a long raised ridge
or keel;
the scales may also have small knobs and tiny pits. The skin and outer
covering of
the horny scales are shed periodically and usually in one piece, including
the hard,
transparent covering of the eye known as the spectacle (snakes lack movable
eyelids, and the spectacle protects the constantly open eyes). The frequency
of
shedding varies with different species and within species, according to
the size and
age of the individual. Young, rapidly growing snakes shed their skins more
frequently than the slow-growing adults. In some species the skin is shed
about
every 20 days; in others, only once a year.
Structure and Function
The long, slender body of the snake contains no fewer than 100 vertebrae
and
sometimes more than 300. Except for the first two, which connect with the
head,
each vertebra is connected to a pair of ribs. The skeleton is light and
provides great
freedom of movement. The skull in particular is loosely built and can be
stretched in
several directions, permitting the snake to swallow large meals in relation
to the
size of the head and body. This mobility can be seen especially in the
two bones of
the lower jaw, which are attached to the skull by a short, movable bone
and which
are united at the front end merely by an elastic ligament. Both jaws have
a large
number of sharp, needlelike teeth, all curved toward the rear of the mouth.
The teeth
are arranged in six rows, two rows on each side of the upper jaw and one
on each
side of the lower jaw. Except in venomous species, the teeth are solid
and are
replaced periodically. When the snake catches its prey, the recurved teeth
enable it
to retain a firm hold. The prey is killed quickly and swallowed by alternate
movement
of the rows of teeth, which work to pull the food into the mouth. As the
food passes
through the mouth, it is covered with saliva; contrary to popular belief,
snakes do not
cover their food with saliva before taking it into the mouth. Most meals
are
swallowed easily and quickly, but a large meal may take several hours.
The big
pythons can eat animals that weigh up to about 68 kg (150 lb), but swallowing
such
a meal is a difficult process.
Venomous snakes have two hollow, hypodermiclike teeth, known as fangs,
in the
front of the upper jaw that, like the other teeth, are replaced periodically.
The new
fang moves into place before the old one is lost, so that, for a short
time, the snake
may have two fangs on each side of the jaw. In the solenoglyphs, a large
group of
snakes that includes the vipers, the fang-bearing bone is attached to the
skull in
such a way that the fangs can be moved back up against the roof of the
mouth
when not in use. The cobras and coral snakes represent another large group
of
snakes, the proteroglyphs, in which the fangs are not movable but are constantly
erect. The fangs of both groups are connected by the venom duct to the
two venom
glands, which are modified salivary glands situated on each side of the
head behind
the eye.
The snake must bite to inject its venom; no snake has a stinger in its
tail. Snakes
can bite at any time and from any position, including underwater. They
normally
strike out from a defensive coil, projecting the head and front part of
the body at the
victim. As soon as the head leaves the coil, the mouth is wide open. The
solenoglyphs stab the erect fangs into the target and usually withdraw
them
immediately, either with or without a biting movement. The proteroglyphs
usually
bite, and the bite may be retained for several seconds. Three species of
snake can
spit or eject the venom in a fine spray, which is aimed at the eyes of
an enemy and
projected for distances up to 2.4 m (8 ft). If the venom gets into the
eyes, it may
cause blindness. The spitting is used only in defense and never to obtain
food.
Snake venom damages various tissues of the body, immobilizing or killing
the
snake?s prey. Venoms are usually classified in two major categories, neurotoxic,
or
nerve-affecting, and hemotoxic, or blood-affecting. Because of local damage
to
circulatory cells, hemotoxic venom spreads more slowly and has a slower
action
than the neurotoxins.
Although snakes lack a true voice, they often hiss loudly. The bull snakes
and their
relatives, for example, have a small flap in front of the opening of the
windpipe that
enables them to produce an extremely loud hiss.
Vision is well developed in most snakes, but many burrowing snakes are
virtually
blind. Snakes have a strong sense of smell, which is relied on to a large
extent in
hunting food. They also have an extra chemical sense that is widely used
during
most activities. The tongue flicks out, picking up odors and carrying them
to the roof
of the mouth into contact with a sensory receptor called Jacobson?s organ.
The
tongue also receives touch stimuli. Snakes are deaf to airborne sounds.
Thus a
rattlesnake does not hear the sound of another snake?s rattle, nor does
the cobra
hear, as it is believed, the snake-charmer?s flute. They can, however,
feel vibrations
through the ground or whatever they are resting on.
The pit vipers, as well as some boas and pythons, have another kind of
sense
organ, a heat receptor that can detect small differences in temperature.
In the pit
vipers the heat receptor is located in facial pits, between the eye and
nostril on each
side of the face; in the boas and pythons the receptor is located in the
labial pits.
By means of these receptors the snakes can locate and seize warm-blooded
prey
at night.
Locomotion
Most snakes lack external limbs, although a few have vestigial hind legs,
the tips of
which are visible at the base of the tail in large boas and pythons. A
surprising
characteristic of snakes is their ability to move rapidly without legs.
Four different
types of locomotion are used by snakes in getting from place to place,
although not
all snakes can use all four methods. The most commonly used method is the
simple, undulating crawl, which is called the serpentine method. In this
type of
locomotion, the snake pushes against the ground on the back side of each
curve or
undulation and flows smoothly forward. All snakes use the serpentine method
for
maximum speed. Snakes move slower than an adult human can run; the fastest
recorded speed achieved by any snake is about 13 km/hr (8 mph), but few
can go
that fast. In another method called caterpillar, used only by the heavier-bodied
snakes, the skin of the ventral surface is moved forward and backward by
strong
muscles, and the broad belly scales grip the ground, moving the snake forward
in a
straight line. This method has led to the mistaken belief that snakes walk
on their
ribs; actually the ribs do not move forward and backward in any of the
four types of
movement. Several desert-dwelling species use a special type of locomotion,
called
sidewinding, to move on loose sand. In this method the snake rolls its
body
sidewise along the ground in a looping motion. The fourth method is known
as
concertina, because the body is alternately stretched out and pulled together
as the
snake moves from one anchor point to another. The concertina is used in
crossing
smooth surfaces and in climbing. Except for sidewinding, any of the methods
may
be used in climbing, but snakes swim only by means of the serpentine method.
Some species of the Colubridae family, such as garters, and king snakes,
in East
Asia and New Guinea reportedly fly. These snakes do not actually fly, but
they can
drop or hurl themselves from fairly high trees and fall or even partly
glide to the
ground without injury. It was once mistakenly believed that the mud snake
could
take its tail in its mouth and roll along like a hoop, which gave rise
to its popular
name, the hoop snake.
Behavior
Snakes have a well-developed nervous system and an intelligence intermediate
between that of fish and mammals. They adapt well to captivity, and many
become
quite tame. The members of many species are nervous and excitable when
first
caught but usually respond quickly to gentle care. When disturbed in the
wild, some
snakes, such as the hognose, go through elaborate bluffing performances;
harmless
snakes can even appear more dangerous than venomous ones.
Many snakes vibrate their tails back and forth when excited, and if they
happen to
be in dry grass or leaves, a whirring noise may be heard. The rattlesnake?s
horny
rattles, as an extreme example, are vibrated at an average rate of 50 times
per
second. The rattling sound, like the hiss, is a warning to enemies; it
is not a lure or
a mating sound or a signal to other rattlesnakes. Contrary to popular opinion,
the
number of rattles does not indicate the age of the snake. A new segment
or rattle is
added every time the snake sheds its skin, which occurs from one to five
times a
year, and many segments are broken off from time to time.
All snakes are carnivorous, eating a variety of animal life, from insects,
spiders, and
snails to frogs, mice, and rats. Oddly, a number of snakes eat only other
kinds of
snakes.
Reproduction
Snakes reproduce either by laying eggs, which the female usually abandons
in a
secluded spot, or by bringing forth living young from eggs hatched within
the body,
which are also abandoned. Eggs and young may number as many as 100 at one
time, but much smaller clutches are usual. Parental care is entirely absent
so far as
the young are concerned, but a few species guard the eggs during incubation,
and
the female python actually broods them. Growth is fairly rapid, and the
young reach
maturity in from one to five years. The greatest age known for any snake
is just
under 30 years, attained by both the anaconda and the black-lipped cobra.
Habitat
Snakes are found in all the warmer parts of the world, except Ireland,
New Zealand,
and a number of oceanic islands. They occur as far north as the Arctic
Circle, but
the variety decreases from the Tropics poleward.
Importance
Snakes are the most recent group of reptiles to appear, probably evolving
from
lizards or lizardlike animals about 100 million years ago, toward the end
of the
Mesozoic era. Human encroachment upon their habitats, as well as the belief
that
they are dangerous and must be exterminated, threatens to diminish their
numbers.
Through their carnivorous food habits, however, snakes often play an important
part
in preserving an ecological balance, particularly in controlling such pests
as
rodents, which tend to multiply at a rapid rate. They also help to keep
down a
number of species injurious to agriculture.
The snake has been held in mythical or religious awe by various cultures
throughout
history. The ancient Egyptians worshipped the asp as a symbol of fertility;
the
Greeks and Romans associated the Aesculapian snake with the god of healing
and
medicine; and the serpent figures prominently in the Bible. Among some
American
Indian tribes, the snake has been associated with rain and fertility. The
cobra has
been highly regarded in India, and snakes play a role in the practice of
voodoo in the
West Indies, as well as in snake-handling cults in the U.S. The serpent
also
appears, usually as a symbol of evil, throughout art and literature.
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