REPTILE, common name for any animal of the vertebrate class Reptilia. Millions
of
years ago, during the Mesozoic era, dinosaurs and other reptiles were the
most
numerous vertebrate animals on the earth. Reptiles have since been reduced
from
17 recognized orders to just four:
? Squamata, with about 3000 species of lizards, 2500 of snakes, and 140
of worm
lizards (amphisbaenians);
? Chelonia, or Testudines, comprised of about 250 species of turtles and
tortoises;
? Crocodylia, including 21 species of crocodiles, alligators, gavials,
and caimans;
? Rhynchocephalia, which flourished during the Jurassic period but today
is
represented only by the tuatara, which lives on several islands off the
coast of New
Zealand. Like most classes of vertebrates (animals that have a backbone)
reptiles
vary in size. The increasingly rare Indian gavial may reach a length of
9 m (30 ft) and
weigh 540 kg (1200 lb). The gecko lizard, by contrast, is typically 3 cm
(1.2 in) long.
Reptiles also include a fascinating range in behavior and appearance; yet,
from the
timid box turtle to the savage Komodo dragon, all share certain traits
that set them
apart as reptiles.
Physical Characteristics
All reptiles have scaly, thick, waterproof skin that is dry to the touch.
Turtles and
tortoises have wide, flat ribs over which is laid the "shell" of horny
or leathery scales
that serves as protective armor. Snakes and many other species molt: As
they
grow, they periodically shed the outer skin, revealing a new layer that
has grown
under the old. Turtles and tortoises gain new shells in much the same way.
Internal Structure
All reptiles, even aquatic species, breathe through lungs. The serpentlike
eel may
look like a reptile, but it is classified as a fish because it breathes
through gills
instead of lungs. In most snakes and some lizards only one lung is functional;
in
crocodiles and other reptiles both lungs are equally developed. The thorax
and
abdomen in reptiles are not separated by a diaphragm, and breathing is
accomplished by muscles of the body wall. The reptilian heart generally
consists of
three chambers, two auricles and one ventricle, in contrast to the mammalian
heart,
which has two auricles and two ventricles. In the crocodilian heart, however,
the
ventricle is almost completely divided into two chambers by a septum, or
partition.
Blood is pumped from the heart to the rest of the reptile?s body through
arteries.
Venous blood returns to the heart from the tail and hind limbs through
the kidneys
by way of a renal portal (vein-capillary); blood from the abdominal region
returns by a
portal system passing through the liver. Carnivorous reptiles frequently
have fairly
simple digestive systems; those that are herbivorous have a more complicated
arrangement. All have an intestine for excretion; the urinary bladder is
present only
in turtles and lizards.
Body Temperature
Reptiles are commonly described as cold-blooded, although the term can
be
misleading. When active, some reptiles maintain their bodies at a higher
temperature than do most mammals. The important difference in temperature
physiology is that reptiles rely on external sources of heat, whereas birds
(which
may be descended from ancient reptiles) and mammals generate internal heat.
Reptiles regulate their body temperatures by taking advantage of sources
of outside
warmth, such as direct sunlight, warm surfaces, and the earth itself. Often
the body
temperature of a reptile is well above that of the surrounding air. Only
when the
animal is inactive or dormant is body temperature approximately the same
as that of
the environment.
Eating Habits
Crocodiles, snakes, and lizards are meat-eaters, as is the tuatara. Turtles
and
tortoises can be categorized as omnivorous; those that live on land generally
eat
vegetable matter, but aquatic species are largely carnivorous.
Reproduction
Most reptiles reproduce sexually, and most mate in the spring. Female turtles,
crocodilians, and the tuatara are primitively oviparous?that is, they lay
eggs. Most
snakes and lizards lay eggs as well. Other reptiles, such as the anaconda,
are
viviparous?that is, they give birth to living young; and some, such as
vipers, are
ovoviparous?that is, they give birth to living young from eggs hatched
inside the
mother?s body. Some large sea turtles may lay as many as 200 eggs in a
clutch;
the tiny geckos generally lay two eggs at a time. Reptile eggs have shells
and
contain two embryonic membranes: a protective amnion, or egg sac, which
prevents
the egg from drying out, and an allantois, or vascular fetal membrane that
serves as
a bladder and a kind of lung. In many species the eggs are left to develop
by
themselves in a sheltered area, where they may draw heat from the sun or
the
overall environment and be protected from predators. Many, however, are
lost to
birds, mammals, and other reptiles.
Habitat
Reptiles inhabit most parts of the world, but because of their need to
obtain warmth
from the environment, they do not live or develop in colder regions. Most
are found
between 40°N and 40°S lat. In temperate regions they may hibernate
during the
winter, and in hot climates they frequently estivate?lie motionless, in
a state of
torpor, during periods of intense heat. Thus, reptiles in the Tropics generally
are
more active at night than during the day, when they tend to lie in the
shade.
Water-dwelling turtles and tortoises are adapted to the ambient temperature
of the
environment where they live, but none are found in the Arctic or Antarctic
regions.
Evolution
The earliest reptiles developed from amphibians during the Upper Carboniferous
period of the Paleozoic era about 300 million years ago. Many forms evolved
and
flourished in the Mesozoic era, the Age of Reptiles. The best-known orders
of
extinct reptiles are as follows:
Ichthyosauria
These large marine reptiles had bodies that resembled those of present-day
fish.
Some species were more than 4 m (13 ft) long.
Sauropterygia
This order included long-necked aquatic forms that had large bodies and
limbs
adapted for moving through the water.
Saurischia and Ornithischia
These orders included the reptiles commonly known as dinosaurs. First found
in
Triassic strata, they became extinct at the close of the Mesozoic era.
Pterosauria
This large, diverse group of flying reptiles is related to the dinosaurs.
Their wings
were thin membranes attached to the body by the very long fourth finger
of each
arm.
Therapsida
Commonly called "mammallike reptiles," these were not reptiles at all,
but
relatives of mammals that split from Reptilia by the end of the Carboniferous
period.
Recent evidence indicates that at least some extinct reptiles were warm-blooded.
In particular, pterosaurs (commonly called pterodactyls), the flying reptiles,
had a
thick fur coat, indicating the need to insulate body warmth. It is also
likely that at
least some dinosaurs, with their rapid growth rate and erect posture, were
warm-blooded. Because of this some zoologists have proposed reclassifying
these fossil animals out of the class Reptilia.
Of the almost 6000 species of reptiles still in existence, about 400 are
endangered, most because of human activity. Turtles have been killed for
their
meat, alligators for their hides, and many people regard snakes, in particular,
as
dangerous serpents to be eradicated. Probably the most significant problem,
however, is the encroachment by humans upon traditional reptile habitats.
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